姓　　名：孙青原
学　　科：发育生物学
电话/传真：+86-/
电子邮件：sunqy@ioz.ac.cn, sunqy1@yahoo.com
通讯地址：北京市朝阳区北辰西路1号院5号
中国科学院动物研究所 干细胞与生殖生物学国家重点实验室 100101
更多信息：受精生物学研究组

简历介绍：
　　孙青原，男，1964年12月出生于山东省招远市。中国科学院动物研究所特聘研究员，中国科学院大学岗位教授，国家****基金获得者，“973”项目首席科学家，新世纪百千****才工程国家级人选。1994年毕业于东北农业大学，获动物胚胎学理学博士学位。1996年-2001年在中国科学院动物研究所、以色列巴伊兰大学、美国密苏里大学、日本农林水产省畜产试验场、日本农业生物资源研究所等从事博士后或合作研究。1996年和1998年分别被晋升为副研究员、研究员。
　　目前兼任中国动物学会副理事长、生殖生物学分会理事长、中国解剖学会发育生物学分会副主任委员等。曾担任计划生育生殖生物学国家重点实验室主任、中国动物学会细胞与分子显微技术分会理事长、第六届国务院学位委员会生物学组成员等。长期从事卵子发育、成熟、受精、早期胚胎发育、动物克隆及生殖发育的表观遗传学研究，在Science、Cell、PNAS、Nat Comm、PLoS Genet、Development、EHP、J Cell Sci、JBC、MBC、Hum Reprod、Fertil Steril、Biol Reprod 等杂志发表SCI论文332篇，受邀为领域知名期刊如Mol Aspect Med、Hum Reprod Update、Autophagy、Semin Cell Develop Biol、Mol Endocinol、Cell Cycle、Mol Hum Reprod、Biol Reprod等撰写综述。2014、2015年入选爱思唯尔（Elsevier）中国高被引****榜单（生化、遗传与分子生物学领域）。近5年来担任Biology of Reproduction、Molecular Human Reproduction、Molecular Reproduction and Development、Scientific Reports、Cell Cycle、Reproduction Fertility and Development、Journal of Genetics and Genomics、Journal of Reproduction and Development等15个SCI期刊副主编或编委。曾获全国优秀博士后、中国科学院青年科学家奖、中国科学院****、国家留学回国人员成就奖、中国青年科技奖、“十一五”国家科技计划执行突出贡献奖等荣誉和国家自然科学二等奖1项、省部级一等奖3项、全国妇幼健康科技奖自然科学一等奖等。
杂志编委：
　　《Biology of Reproduction》（SCI）编委会委员(综述委员会) (2007-2009)；编委会委员(审稿编辑委员会) (2009—现在)
　　《Cell Cycle》（SCI）编委员会委员(2007-现在)
　　《Molecular Reproduction and Development》（SCI）副主编(2007-2008) （2014—现在）
　　《Reproduction, Fertility and Development》（SCI）执行副主编 (2009-2016)
　　《Journal of Reproduction and Development》（SCI）编委员会委员(2004-现在)
　　《Asian Journal of Andrology》（SCI）编委员会委员(2004—2015)
　　《Progress in Natural Science》（SCI）编委员会委员(2004--2009)
　　《Prog Biophys Biochem》（SCI）编委员会委员(2010—现在)
　　《Current Zoology》副主编（SCI）（2009—现在）
　　《The Scientific World Journal》编委员会委员（2010—现在）
　　《PLoS ONE》Editor （2012-现在）
　　《Journal of Genetics and Genomics》编委会委员（2014—2016）、副主编（2017-2019）
　　《Peer J》编委会委员
　　《动物学报》编委员会委员(1999--2008)
　　《动物学杂志》编委员会委员(1999—现在)
　　《生殖医学杂志》编委员会委员(2002—现在)
　　《细胞生物学杂志》编委员会委员(2007—2009)
　　《中国细胞生物学报》编委员会委员(2010-现在)
　　《遗传》副主编（2010—2013）
　　《生殖与避孕》副主编（2012-现在）
　　《Scientific Reprots》（SCI）编委员会委员(2015—现在)
　　《Molecular Human Reproduction》副主编(2017—)
学历及培训经历:
　　1999 年 10 月－2001 年 8 月 University of Missouri-Columbia, USA 博士后 (Research associate)
　　1996 年 10 月－1998 年 4 月 Bar-Ilan University, Israel 博士后
　　1994 年 8 月－1996 年 8 月 中国科学院动物研究所生殖生物学国家重点实验室 博士后
　　1991 年 8 月－1994 年 6 月 东北农业大学生物工程系 博士研究生
　　1985 年 7 月－1988 年 6 月 东北农业大学生物工程系 硕士研究生
　　1981 年 9 月－1985 年 6 月 山东农业大学畜牧兽医系 本科生
工作经历:
　　1996年9月-现在 中国科学院动物研究所副研究员（1996）、研究员（1998），博士生导师（2000），生殖生物学中心主任（2002-2006），计划生育生殖生物学国家重点实验室常务副主任（2002-2006）、主任（2006-2011）
　　2009年4月-2005年7月 University of Missouri-Columbia，USA 高级研究****
　　2008 年 4 月－2008 年 6 月 National Institute of Agrobiological Sciences, Japan, Invited Short-Term JAPS Fellow
　　2005 年 5 月－2005 年 8 月 University of Missouri-Columbia，USA 高级研究****
　　2003 年 3 月－2003 年 4 月 National Institute of Agrobiological Sciences，Japan, Invited Short-Term JAPS Fellow
　　1996 年 1 月－1996 年 4 月 National Institute of Animal Industry, Japan STA Fellow
　　1988 年 7 月－1991 年 7 月 山东农业大学畜牧兽医系 助教
研究领域：
卵母细胞减数分裂和受精的信号转导
卵母细胞减数分裂纺锤体组装、纺锤体检验点和染色体分离调节
卵子和早期胚胎发育（包括动物克隆）的表观遗传
社会任职：
获奖及荣誉：
承担科研项目情况：
国家自然科学基金面上项目：urvivin在小鼠卵母细胞减数分裂成熟中的作用及机制研究（**），2013-2016
国家自然科学基金面上项目：利用小鼠模型研究PCOS对小鼠卵母细胞及后代生殖细胞DNA甲基化印迹影响（**），2015-2018
国家自然科学基金重点项目：蛋白磷酸酶2A类家族在小鼠卵子发生和成熟中的作用及机制研究（**），2016-2020
国家重大科学研究计划项目：雌性生殖细胞减数分裂的分子基础（2012CB944400），2012-2016
代表论著：
Zhang T, Zhou Y, Li L, Wang ZB, Shen W, Schatten H, Sun QY*. CenpH regulates meiotic G2/M transition by modulating the APC/CCdh1-cyclin B1 pathway in oocytes. Development. 2016 Dec 19. pii: dev.141135. [Epub ahead of print]
Wang HH, Cui Q, Zhang T, Guo L, Dong MZ, Hou Y, Wang ZB, Shen W, Ma JY, Sun QY*. Removal of mouse ovary fat pad affects sex hormones, folliculogenesis and fertility. J Endocrinol. 2017 Feb;232(2):155-164.
Hu MW, Meng TG, Jiang ZZ, Dong MZ, Schatten H, Xu X*, Wang ZB*, Sun QY*. Protein Phosphatase 6 Protects Prophase I-Arrested Oocytes by Safeguarding Genomic Integrity. PLoS Genet. 2016 Dec 8;12(12):e**. doi: 10.1371/journal.pgen.**.
Liu K, Zhao Q, Liu P, Cao J, Gong J, Wang C, Wang W, Li X, Sun H, Zhang C, Li Y, Jiang M, Zhu S,Sun Q, Jiao J, Hu B, Zhao X, Li W, Chen Q, Zhou Q, Zhao T. ATG3-dependent autophagy mediates mitochondrial homeostasis in pluripotency acquirement and maintenance. Autophagy. 2016 Nov;12(11):2000-2008.
Qi ST, Ma JY, Wang ZB, Guo L, Hou Y, Sun QY*. N6-methyladenosine seqencing highlights the involvement of mRNA methylation in oocyte meiotic maturation and embryo development by regulating translation in Xenopus laevis. J Biol Chem. 2016 Oct 28;291(44):23020-23026.
Ma XS, Lin F, Wang ZW, Hu MW, Huang L, Meng TG, Jiang ZZ, Schatten H, Wang ZB*, Sun QY*. Geminin deletion in mouse oocytes results in impaired embryo development and reduced fertility. Mol Biol Cell. 2016, 27(5):768-75.
Wang ZB, Ma XS, Hu MW, Jiang ZZ, Meng TG, Dong MZ, Fan LH, Ouyang YC, Snapper SB, Schatten H, Sun QY*. Oocyte-specific deletion of N-WASP does not affect oocyte polarity, but causes failure of meiosis II completion. Mol Hum Reprod. 2016 Sep;22(9):613-21.
Yu C, Ji SY, Sha QQ, Wang ZW, Sun QY, Fan HY. BTG4 is a meiotic cell cycle-coupled maternal-zygotic tansition licensing factor in oocytes. Nature Structural and Molecular Biology 2016 May;23(5):387-94.
Jiang ZZ, Hu MW, Ma XS, Schatten H, Fan HY, Wang ZB, Sun QY*. LKB1 acts as a critical gatekeeper of ovarian primordial follicle pool. Oncotarget. 2016, 7(5):5738-53.
Zhang T, Zhou Y, Li L, Wang HH, Ma XS, Qian WP, Shen W, Schatten H,Sun QY*. SIRT1, 2, 3 protect mouse oocytes from postovulatory aging. Aging (Albany NY). 2016, 8(4):685-96.
Ma XS, Lin F, Wang ZW, Hu MW, Huang L, Meng TG, Jiang ZZ, Schatten H, Wang ZB, Sun QY*. Geminin Deletion in Mouse Oocytes Results in Impaired Embryo Development and Reduced Fertility. Mol Biol Cell. 2016 Jan 13. pii: mbc.E15-06-0346. [Epub ahead of print]
Yu C, Ji SY, Sha QQ, Sun QY, Fan HY. CRL4-DCAF1 ubiquitin E3 ligase directs protein phosphatase 2A degradation to control oocyte meiotic maturation. Nat Communications. 2015 Aug 18;6:8017.
Hu MW, Wang ZB, Teng Y, Jiang ZZ, Ma XS, Hou N, Cheng X, Schatten H, Xu X*, Yang X*, Sun QY*. Loss of protein phosphatase 6 in oocytes causes failure of meiosis II exit and impaired female fertility. J Cell Sci. 2015 Oct 15;128(20):3769-80. (Commented in this issue, cover paper)
Wei Y, Yang CR, Wei YP, Ge ZJ, Zhao ZA, Zhang B, Hou Y, Schatten H, Sun QY*. Enriched Environment-Induced Maternal Weight Loss Reprograms Metabolic Gene Expression in Mouse Offspring. J Biol Chem. 2015, 20;290(8):4604-19.
Wei YC，Schatten H, Sun QY*. Environmental epigenetic inheritance through gametes and implications for human reproduction. Hum Reprod Update 2015 Mar-Apr;21(2):194-208
Wei Y, Yang CR, Wei YP, Zhao ZA, Hou Y, Schatten H, Sun QY*. Paternally induced transgenerational inheritance of susceptibility to diabetes in mammals.Proc Natl Acad Sci USA （PNAS） 111(5):1873-1878, 2014

Yu XJ, Yi Z, Gao Z, Qin DD, Zhai Y, Chen X, Ou-Yang YC, Wang ZB, Zheng P, Zhu MS, Wang H, Sun QY Dean J, Li L. The subcortical maternal complex controls symmetric division of mouse zygotes by regulating F-actin dynamics. Nature Communications 2014 Sep 11;5:4887.
Ge ZJ, Luo SM, Lin F, Liang QX, Huang L, Wei YC, Hou Y, Han ZM, Schatten H, Sun QY*. DNA methylation in oocytes and liver of female mice and their offspring: Effects of high-fat-diet-induced obesity. Environmental Health Perspectives 122(2):159-164, 2014.

Jiang ZZ, Hu MW, Wang ZB, Huang L, Lin F, Qi ST, Ouyang YC, Fan HY, Schatten H, Mak TW, Sun QY*. Survivin is essential for fertile egg production and female fertility in mice. Cell Death Dis. 2014 Mar 27;5:e1154. doi: 10.1038/cddis.2014.126.

Ge ZJ, Zhang CL, Schatten H, Sun QY*. Maternal diabetes mellitus and the origin of non-communicable diseases in offspring: the role of epigenetics. Biol Reprod. 2014 May 14. pii: biolreprod.114.118141. [Epub ahead of print]

Lin F, Ma XS, Wang ZB, Wang ZW, Luo YB, Huang L, Jiang ZZ, Hu MW, Schatten H, Sun QY*. Different fates of oocytes with DNA double-strand breaks in vitro and in vivo. Cell Cycle 2014, 13(17): in press.

Hu MW, Wang ZB, Jiang ZZ, Qi ST, Huang L, Liang QX, Schatten H, Sun QY*. Scaffold Subunit Aalpha of PP2A Is Essential for Female Meiosis and Fertility. Biol Reprod. 2014 Jun 4. pii: biolreprod.114.120220. [Epub ahead of print]

Qiao J, Wang ZB, Feng HL, Miao YL, Wang Q, Yu Y, Wei YC, Yan J, Wang WH, Shen W, Sun SC, Schatten H, Sun QY*. The root of reduced fertility in aged women and possible therapentic options: Current status and future perspects. Molecular Aspects of Medicine 2013 Jun 21. doi:pii: S0098-2997(13)00041-1. 10.1016/j.mam.2013.06.001. [Epub ahead of print], 2014

Yu C, Zhang YL, Pan WW, Li XM, Wang ZW, Ge ZJ, Zhou JJ, Cang Y, Tong C, Sun QY*, Fan HY*. CRL4 complex regulates mammalian oocyte survival and reprogramming by activation of TET proteins. Science 342:1518-21,2013. （Highlighted by Biol Reprod, 2014）

Wang ZB, Jiang ZZ, Zhang QH, Hu MW, Huang L, Ou XH, Guo L, Ouyang YC, Hou Y, Brakebusch C, Schatten H, Sun QY*. Specific deletion of Cdc42 does not affect meiotic spindle organization/migration and homologous chromosome segregation, but disrupts polarity establishment and cytokinesis in mouse oocyte. Molecular Biology of the Cell 24: 3832-3841, 2013.

Luo SM, Ge ZJ, Wang ZW, Jiang ZZ, Wang ZB, Ouyang YC, Hou Y, Schatten h, Sun QY*. Unique insights into maternal mitochondrial inheritance in mice. Proc Natl Acad Sci USA （PNAS）110(32):13038-13043, 2013.

Wang ZW, Ma XS, Ma JY, Luo YB, Lin F, Wang ZB, Schatten H, Fan Hy, Sun QY*. Laser Microbeam-Induced DNA Damage Inhibits Cell Division in Fertilized Eggs and Early Embryos. Cell Cycle 12(20):3336-3344,2013.

Luo SM, Sun QY*. Autophagy Is Not Involved in the Degradation of Sperm Mitochondria after Fertilization in Mice. Autophagy 9(12):2156-2157,2013.

Xia P, Wang S, Du Y, Zhao Z, Shi L, Sun L, Huang G, Ye B, Li C, Dai Z, Hou N, Cheng X, Sun QY, Li L, Yang X, Fan ZWASH inhibits autophagy through suppression of Beclin 1 ubiquitination. EMBO Journal. 32(20):2685-2696,2013.

Ma JY, Li M, Luo YB, Song S, Tian D, Yang J, Zhang B, Hou Y, Schatten H, Liu Z, Sun QY*. Maternal factors required for oocyte developmental competence in mice: Transcriptome analysis of non-surrounded nculeolus (NSN) and surrounded nucleolus (SN) oocytes. Cell Cycle. 12:1928-1938, 2013.

Qi ST, Wang ZB, Ouyang YC, Zhang QH, Hu MW, Huang X, Ge ZJ, Guo L, Wang YP, Hou Y, Schatten H, Sun QY* Overexpression of SETβ, a protein localizing to centromeres, causes precocious separation of chromatids during the first meiosis of mouse oocyte. Journal of Cell Science 126, 1595-1603, 2013.

Luo YB, Ma JY, Zhang QH, Lin F, Wang ZW, Huang L, Schatten H, Sun QY*. MBTD1 is associated with Pr-Set7 to stabilize H4K20me1 in mouse oocyte meiotic maturation. Cell Cycle. 12(7): 1142-1150, 2013.

Ma JY, Ouyang YC, Luo YB, Wang ZB, Hou Y, Han ZM, Liu ZH, Schatten H, Sun QY*Cyclin O regulates germinal vesicle breakdown in mouse oocytes. Biology of Reproduction, 88(5):110, 1-9, 2013.

Ge ZJ, Liang XW, Guo L, Liang QX, Luo SM, Wang YP, Wei YC, Han ZM, Schatten H, Sun QY* Maternal diabetes causes alterations of DNA imprinting in murine oocytes. Biology of Reproduction, 88(5):117, 1-9, 2013.

（Reported by Science Daily, Science News, Medical News Today, Medical Express, MedSci）

Ma JY, Ouyang YC, Wang ZW, Wang ZB, Jiang ZZ, Luo SM, Hou Y, Liu ZH, Sun QY* The effects of DNA double-strand breaks on mouse oocyte meiotic maturation. Cell Cycle 12(8): 1928-1938, 2013.

Yang SW, Gao C, Chen L, Song YL, Zhu JL, Qi ST, Jiang ZZ, Wang ZW, Lin F, Huang H, Xing FQ, Sun QY* Nek9 regulates spindle organization and cell cycle progression during mouse oocyte meiosis and its location in early embryo mitosis. Cell Cycle 16;11(23):4366-77，2012.

Ma JY, Liang XW, Schatten H, Sun QY* Active DNA demethylation in mammalian preimplantation embryos: new insights and new perspectives. Molecular Human Reproduction. 18(7):333-40，2012.（Cover paper）

Ou XH, Li S, Wang ZB, Li M, Quan S, Xing F, Chao SB, Chen ZJ, Liang XW, Hou Y, Schatten H, Sun QY*. Maternal insulin resistance causes oxidative stress and mitochondrial dysfunction in mouse oocytes. Human Reproduction 27(7):2130-45,2012.

Li S, Ou XH, Wei L, Wang ZB, Zhang QH, Ouyang YC, Hou Y, Schatten H, Sun QY*. Septin 7 is required for orderly meiosis in mouse oocytes. Cell Cycle. 11(17):3211-8，2012.

Chen L, Chao SB, Wang ZB, Qi ST, Zhu XL, Yang SW, Yang CR, Zhang QH, Ouyang YC, Hou Y, Schatten H,Sun QY* Checkpoint kinase 1 is essential for meiotic cell cycle regulation in mouse oocytes. Cell Cycle 15;11(10):1948-55, 2012.

Chao SB, Guo l, Ou XH, Luo SM, Wang ZB, Schatetn H, Gao l, Sun QY* Heated spermatozoa: effects on embryonic development and epigenetics. Human Reproduction 27(4):1016-24,2012.

Zhang QH, Qi ST, Wang ZB, Yang CR, Wei YC, Chen L, OuYang YC, Hou Y, Schatetn H, Sun QY* Localization and function of the Ska complex during mouse oocyte meiotic maturation. Cell Cycle 11(5):909-16, 2012.

Zhu XL, Qi ST, Liu J, Chen L, Zhang C, Yang SW, OuYang YC, Hou Y, Schatten H, Sun QY* Synaptotagmin1 is required for spindle stability and metaphase-anaphase transition in mouse oocytes. Cell Cycle 11(4):818-26, 2012.

Wang ZB, Schatten H, Sun QY* Why is chromosome segregation rrror in oocytes increased with maternal aging? Physiology 26:314-325, 2011.（cover paper）

Sun QY* Ndc80 complex: New evidence for the existence of spindle assembly checkpoint in mammalian oocyte meiosis. Cell Cycle 15;10(6):877-8, 2011.

Liang XW, Ge ZJ, Guo L, Luo SM, Han ZM, Schatten H, Sun QY* Effect of postovulatory oocyte aging on DNA methylation imprinting acquisition in offspring oocytes. Fertility and Sterility.96: 1479-1484, 2011.

Zhang CH, Wang ZB, Quan S, Huang X, Tong JS, Ma JY, Guo L, Wei YC, Ouyang YC, Hou Y, Xing FQ, Sun QY* GM130, a cis-Golgi protein, regulates meiotic spindle assembly and asymmetric division in mouse oocyte. Cell Cycle 10(11):2230-2236, 2011.

(Commented by Sun SC in Cell Cycle, 2011; Liu XJ in Cell Cycle, 2011)

Liang XW, Ge ZJ, Wei L, Guo L, Han ZM, Schatten H, Sun QY* The effects of postovulatory aging of mouse oocytes on methylation and expression of imprinted genes at mid-term gestation. Molecular Human Reproduction 17:562-567, 2011.

Schatten H*, Sun QY* The role of centrosomes in fertilization, cell division and establishment of asymmetry during embryo development. Seminars in Cell & Developmental Biology. 21:174-184, 2010.

Gu L, Wang Q*, Sun QY* Histone modifications during mammalian oocyte maturation: dynamics, regulation, and functions. Cell Cycle 23;9(10)：1942-1950, 2010.

Ou XH, Li S, Xu BZ, Wang ZB, Quan S, Li M, Zhang QH, Ouyang YC, Schatten H, Xing FQ*, Sun QY* p38α MAPK Is a MTOC-Associated Protein Regulating Spindle Assembly, Spindle Length and Accurate Chromosome Segregation during Mouse Oocyte Meiotic Maturation. Cell Cycle 9:4130-4143, 2010. （Faculty of 1000）

Wang ZB, Ou XH, Tong JS, Li S, Wei l, Ouyang YC, Hou Y, Schatten H, Sun QY * The SUMO pathway functions in mouse oocyte maturation. Cell Cycle 9(13):2640-2046, 2010.

Qiao J, Chen Y, Yan LY, Yan J, Liu P, Sun QY. Changes in histone methylation during human oocyte maturation and IVF- or ICSI-derived embryo development. Fertil Steril. 93(5):1628-36, 2010

Zhang QH, Wei L, Tong JS, Qi ST, Li S, Ou XH, Ouyang YC, Hou Y, An LG, Schatten H, Sun QY* Localization and function of mSpindly during mouse oocyte meiotic maturation. Cell Cycle 9(11):2230-2236,2010.

Zhu JL, Lin SL, Li m, Ouyang YC, Hou Y, Schatten H, Sun QY* Septin2 is modified by SUMOylation and required for chromosome congression in mouse oocytes. Cell Cycle 9(8) 1607-1616,2010.

Sha W, Xu BZ, Li M, Liu D, Feng HL, Sun QY* Impact of gonadotrophins on oocyte maturation in vitro: An animal model. Fertility and Sterility 93(5):1650-1661, 2010.

Wei L, Liang XW, Zhang QH, Li M, Yuan J, Li S, Sun SC, Ouyang YC, Schatten H, Sun QY* BubR1 is a spindle assembly checkpoint protein regulating meiotic cell cycle progression of mouse oocyte. Cell Cycle 9(6) 1112-1121, 2010.

（Cover paper, commented by Fan HY. Cell Cycle. 2010 Apr 15;9(8); Jones KT, Holt JE. Cell Cycle. 2010 Mar 15;9(6)）

Ma CH, Yan LY, Qiao J, Sha W, Li L, Chen Y, Sun QY Effects of tumor necrosis factor-alpha on porcine oocyte meiosis progression, spindle organization, and chromosome alignment. Fertil Steril. 93:920-926, 2010.

Sun QY*, Miao YL, Schatten H* Towards a new understanding on the regulation of mammalian oocyte meiosis resumption. Cell Cycle 8(17):2741-7, 2009.

Miao YL, Kikuchi K, Sun QY*, Schatten H* Oocyte aging: implications for developmental potential, and practical significance for assisted reproduction technologies. Human Reproduction Update 15:573-585, 2009.(cover paper)

Yuan J, Sun QY*. Astrin regulates meiotic spindle organization, spindle pole tethering and cell cycle progression in mouse oocytes. Cell Cycle 8(20) 3384-3395, 2009.

Yan LY, Qiao J, Chen Y, Huang J, Liu P, Sun QY Application of 3D- fluorescence in situ hybridization to human preimplantation genetic diagnosis. Fertility and Sterility 92(4):1492-5, 2009.

Sun SC, Wei L, Li M, Lin SL, Xu BZ, Liang XW, Kim NH, Schatten H, Lu SS, Sun QY* Perturbation of survivin expression affects chromosome alignment and spindle checkpoint in mouse oocyte meiotic maturation. Cell Cycle 8(20) 3365-3372, 2009.

Schatten H, Sun QY* The role of centrosomes in mammalian fertilization and its significance for ICSI. Molecular Human Reproduction 15:531-538, 2009.

Li M, Schatten H, Sun QY* Androgen receptor’s destiny in mammalian oocytes: a new hypothesis. Molecular Human Reproduction 15:149-154, 2009.

Sun Y, Kucej M, Fan HY, Yu H, Sun QY, Zou H Separase is recruited to mitotic chromosomes to dissolve sister chromatid cohesion in a polynucleotide-dependent manner. Cell 137:123-132, 2009.

写给考生的话：
　　选择了这一研究方向，就要踏踏实实工作，时间长了就会对研究产生兴趣。相信只要努力工作，前途是光明的。