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基本信息 The basic information
姓名: 郝占庆

学院: 生态环境学院


学历: 博士研究生毕业

学位: 理学博士


职称: 教授

职务:


学科:









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工作经历 Work Experience
1988.7-1996.8 中国科学院沈阳应用生态研究所，助理研究员
1996.9-1999.8 中国科学院沈阳应用生态研究所，副研究员
1999.9-2020.11 中国科学院沈阳应用生态研究所，研究员
1994.9-1995.4 美国俄勒岗州立大学林学系，访问****
1996.11-2002.5 中国科学院长白山森林生态系统开放站，业务副站长
1997.5- 2015.9 中国科学院沈阳应用生态研究所，森林生态研究中心主任
1999.11-2000.5 美国威斯康星大学森林生态与景观实验室，高访
2004.5- 2015.9 中国科学院沈阳应用生态研究所，所长助理
2005.5- 2015.9 中国共产党中国科学院沈阳应用生态研究所委员会，委员
2011.12-2015.11 森林与土壤生态国家重点实验室，副主任、学术委员会委员
2013.5-2015.9 中国科学院沈阳应用生态研究所，科技处长（兼）
2016.4-至今 中国科学院森林生态与管理重点实验室主任
2020.11-至今 西北工业大学生态环境学院，教授








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教育经历 Education Experience
1981.9-1985.7 山西农业大学林学系，获农学学士学位
1985.9-1988.7 中国科学院沈阳应用生态研究所，获理学硕士学位
1997.9-2000.7 中国科学院沈阳应用生态研究所，获理学博士学位







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教育教学 Education And Teaching
自2003年被遴选为中国科学院沈阳应用生态研究所博士生指导教师以来，至今共培养研究生29人，其中博士研究生19人，硕士研究生10人。所培养的博士，大多已成为国内知名大学的教授、博导。2013年获中国科学院“朱李月华优秀教师奖”。
讲授本科生课程《普通生态学》、《植物地理学》、《生态文明与人类未来》等。





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科学研究 Scientific Research
1988年以来一直从事森林生态学、生物多样性等方面的研究工作。在国内率先以长白山温带森林25 ha 大型固定样地为主要平台，系统开展了物种空间分布格局、种间关系、生物多样性维持机制、种子雨及幼苗的时空动态、土壤与凋落物的时空异质性、群落动态等监测和研究，对基于热带森林所发展的物种共存等相关理论给予了验证或完善，相关论文在 Ecology、PNAS、Natural communications、Journal of Ecology、Global Ecology and Biogeography、Ecography、Oecologia、Oikos 等生态学国际主流刊物发表。迄今，已在国内外生态学、林学等主流刊物上发表论文170余篇，总IF 超过320，总被引6717次，H 指数43，i10指数132,其中IF>3 的SCI 论文50 篇。出版《长白山植物多样性及其格局》、《长白山温带森林—阔叶红松林及其次生杨桦林的物种组成及其分布格局(Changbaishan Forest Dynamic Plots--Species Composition and Their Spatial Patterns)》《秦岭皇冠森林动态样地——树种及其分布格局》专著3部，同时参加了其它5部专著的编写。
入职西北工业大学生态环境学院以来，于2019年建立秦岭皇冠25公顷森林样地。目前,已完成地形测绘、植被调查以及土壤理化性质测定等工作，该样地也已正式加入中国森林生物多样性监测网络（CForBio）。秦岭皇冠样地将是西北工业大学生态环境学院科学研究和教育教学的重要平台。
主持科研项目：

1991-1995 中科院“八五”重大项目专题“人类活动对阔叶红松林生态系统多样性的影响”，经费20 万元
1993-1996 国家自然科学基金项目“水源涵养林防护成熟问题的研究”，经费5 万元
1998-2000 国家自然科学基金项目“长白山暗针叶林生态系统生物多样性及其维持机制的研究” 经费14 万元
2001.11-2004.12 中国科学院创新工程重要方向项目“东北地区100 年LUCC 及其生态环境效应研究”课题“100 年来森林资源变化及其生态环 境效应”，经费60 万元
2003.1-2005.12 国家自然科学基金面上项目“长白山暗针叶林苔藓植物生态功能研究”，经费21 万元
2004.4-2008.4 国际生物多样性监测网络计划项目“长白山森林生物多样性格局与动态”，经费20 万元/年
2006.1-2008.12 国家自然科学基金面上项目“长白山阔叶红松林植物多样性共存机制的研究”，经费29 万元
2006.1-2010.12 国家自然科学基金重大项目专题“长白山主要树种对全球变化的响应与适应”，经费18 万元，主持
2007.1-2009.12 中科院知识创新工程重要方向专题“温带森林群落生物多样性格局及维持机制”，经费60 万元
2008.1-2012.12 “十一五”国家科技支撑计划重点项目专题“长白山森林物种资源监测技术与示范”，经费60 万元
2008.1-2010.12 “CForBio and the CTFS-HSBC Climate Partnership project: Monitor the dynamic change of annual carbon flux in the face of climate change in Changbai Mt. plot”，经费5 万美元
2009-2011 国家自然科学基金面上项目：长白山阔叶红松林植物种群扩散、定植过程及其距离效应，经费29 万元
2011-2015 中国科学院知识创新工程重要方向项目：植物功能性状的时空变异与天然林物种共存，150 万元
2012-2017 科技基础性工作专项：《东北温带针阔混交林区生物多样性调查》（2012FY112000），经费210万元2018.1-2022.12国家自然科学基金重点项目，经费291万元





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学术成果 Academic Achievements
近5年代表性论文：
1. Jia SH, Wang XG, Yuan, Z, Lin F, Ye J, Lin G, Hao Z, Bagchi R. (2020). Tree species traits affect which natural enemies drive the Janzen-Connel effect in a temperate forest. Natural Communications 11: 286
2. Yuan Z, Ali A, Ruiz‐Benito P, Jucker T, Mori A, Wang S, Zhang X, Li H, Hao, Z, Wang X. (2020) Above‐and below‐ground biodiversity jointly regulate temperate forest multifunctionality along a local‐scale environmental gradient, Journal of Ecology, 108:2012–2024.
3. Yunyun Wang, Tong Lyu, Nawal Shrestha, Lisha Lyu, Yaoqi Li, Bernhard Schmid, Robert P. Freckleton, Dimitar Dimitrov, Shuguang Liu, Zhanqing Hao, Zhiheng Wang. Drivers of large‐scale geographical variation in sexual systems of woody plants. Global Ecol Biogeogr. 2020; 29: 546– 557
4. Wang YY, LaMontagne JM, Lin F, Yuan ZQ, Ye J, Wang XG, Hao ZQ.(2020).Similarity between seed rain and neighbouring mature tree communities in an old-growth temperate forest. Journal of Forestry Research, 31(6): 2435-2444.
5. Yin QL, Tian TT, Kou M, Liu PL, Wang L, Hao ZQ, Yue M. (2020). The relationships between photosynthesis and stomatal traits on the Loess Plateau. Global ecology and conservation. 23, e01146.
6. Su YJ, Guo QH, Hu TY, Guan HC, Jin SC, An SZ, Chen XL, Guo K, Hao ZQ, Hu YM, Huang YM, Jiang MX, Li JX, Li ZJ, Li XK, Li XW, Liang CZ, Liu RL, Liu Q, Ni HW, Peng SL, Shen ZH, Tang ZY, Tian XJ, Wang XH, Wang RQ, Xie ZQ, Xie YZ, Xu XN, Yang XB, Yang YC, Yu LF, Yue M, Zhang F, Ma KP. (2020). An updated Vegetation Map of China (1:**). Science Bulletin. 65(13): 1125-1136.
7. Fung Tak, Chisholm Ryan A., Anderson-Teixeira Kristina, Bourg Norm, Brochelman Warren Y., Bunyavejchewin Sarayudh, Chang-Yang Chia-Hao, Chitra-Tarak Rutuja, Chuyong George, Condit Richard, Dattaraja Handanakers S., Davies Stuart J., Ewango Corneille E. N., Fewless Gary, Fletcher Christine, Gunatilleke C. V. Savitri, Gunatilleke I. A. U. Nimal, Hao Zhanqing, Hogan J. Aaron, Howe Robert, Hsieh Changfu, Kenfack David, Lin Yiching, Ma Keping, Makana Jean-Remy, McMahon Sean, McShea William J., Mi Xiangcheng, Nathalang Anuttara, Ong Perry S., Parker Geoffrey, Rau E-Ping, Shue Jessica, Su Sheng-Hsin, Sukumar Raman, Sun I-Fang, Suresh Hebbalalu S., Tan Sylvester, ThomasDuncan, Thompson Jill, Valencia Renato, Vallejo Martha I., Wang Xugao, Wang Yunquan, Wijekoon Pushpa, Woif Amy, Yap Sandra and Zimmerman Jess. (2020). Temporal population variability in local forest communities has mixed effects on tree species richness across a latitudinal gradient. Ecology Letters 23: 160-171.
8. Yuan ZQ, Ali A, Sanaei A, Ruiz‐Benito P, Jucker T, Fang L, Bai E, Ye J, Lin F, Fang S, Hao Z, Wang X. (2021). Few large trees, rather than plant diversity and composition, drive the above-ground biomass stock and dynamics of temperate forests in northeast China. Forest Ecology and Management. 481:118698
9. Sanaei A, Ali A, Yuan Z, Liu S, Lin F, Fang S, Ye J, Hao Z, Michel L, Bai E, Wang X. (2021). Context-dependency of tree species diversity, trait composition and stand structural attributes regulate temperate forest multifunctionality. Science of the total environment. In press.
10. 尹秋龙, 贾仕宏, 何春梅, 张乐乐, 成毅聪, 刘润清, 杨治春, 岳明, 郝占庆. 2020. 秦岭皇冠森林动态样地——树种及其分布格局. “中国森林生物多样性监测网络”丛书 (马克平 主编), 中国林业出版社, 北京.
11. Yuan, Z., Ali, A., Wang, S., Wang, X., Lin, F., Wang, Y., Fang, S., Hao, Z*., Loreau, M. & Jiang, L. (2019) Temporal stability of aboveground biomass is governed by species asynchrony in temperate forests. Ecological indicators, 107, 105661.
12. Tak Fung, Ryan A. Chisholm, Kristina Anderson-Teixeira, Norm Bourg, Warren Y. Brockelman, Sarayudh Bunyavejchewin, Chia-Hao Chang-Yang, Rutuja Chitra-Tarak, George Chuyong, Richard Condit, Handanakere S. Dattaraja, Stuart J. Davies, Corneille E.N. Ewango, Gary Fewless, Christine Fletcher, C.V. Savitri Gunatilleke, I.A.U. Nimal Gunatilleke, Zhanqing Hao, J. Aaron Hogan, Robert Howe, Chang-Fu Hsieh, David Kenfack, YiChing Lin, Keping Ma, Jean-Remy Makana, Sean McMahon, William J. McShea, Xiangcheng Mi, Anuttara Nathalang, Perry S. Ong, Geoffrey Parker, E-Ping Rau, Jessica Shue, Shen-Hsin Su, Raman Sukumar, I-Fang Sun, Hebbalulu S. Suresh, Sylvester Tan, Duncan Thomas, Jill Thompson, Renato Valencia, Martha I. Vallejo, Xugao Wang, Yunquan Wang, Pushpa Wijekoon, Amy Wolf, Sandra Yep, & Jess Zimmerman. Temporal population variability in local forest communities has mixed effects on tree species richness across a latitudinal gradient. Ecology Letters (2019)
13. Zikun Mao, Adriana Corrales, Kai Zhu, Zuoqiang Yuan, Fei Lin, Ji Ye, Zhanqing Hao and Xugao Wang. Tree mycorrhizal associations mediate soil fertility effects on forest community structure in a temperate forest. New Phytologist. (2019) 223: 475–486
14. Fang Shuai, Cadotte MarcWilliam, Yuan Zuoqiang, Lin Fei, Ye Ji, Hao Zhanqing and Wang Xugao. Intraspecific trait variation improves the detection of deterministic community assembly processes in early successional forests, but not in late successional forests. Journal of Plant Ecology. (2019)12(4): 593-602
15. Yuan Zuoqiang, Wang Shaopeng, Ali Arshad, Gazol Antonio, Ruiz-Benito Paloma, Wang Xugao, Lin Fei, Ye Ji, Hao Zhanqing*. 2019. Multiple abiotic and biotic pathways shape biomass demographic processes in temperate forests. Ecology. 100(5)，2650
16. Yunyun Wang, Susan J. Mazer, Robert P. Freckleton, Zuoqiang Yuan, Xugao Wang, Yanjun Du, Luxiang Lin, Xihua Wang, Weiguo Sang, Xiaojuan Liu, Zhanqing Hao*. 2019. Testing mechanisms of compensatory fitness of dioecy in a cosexual world. Journal of Vegetation Science, 30(3): 413-426
17. Fang Shuai, Munoz Francois, Ye Ji, Lin Fei, Yuan Zuoqiang, Kuang Xu, Hao Zhanqing and Wang Xugao.Deterministic processes drive functional and phylogenetic temporal changes of woody species in temperate forests in Northeast China.Annals of Forest Science, 2019, 76(2), 42
18. Wang Yunyun, LaMontagne Jalene M., Lin Fei, Yuan Zuoaiqng, Ye Ji, Wang Xugao and Hao Zhanqing*. Similarity between seed rain and neighbouring mature tree communities in an old-growth temperate forest.Journal of forestry research, (2019)
19. 毛子昆, 郝占庆, 原作强, 蔺菲, 叶吉, 匡旭, 王绪高. 物种聚集分布与种间关系的多度不对称性. 中国科学: 生命科学, 2019, 49.
20. 王国杰, 蔺菲, 胡家瑞, 原作强, 郝占庆, 王绪高. 氮和土壤微生物对水曲柳幼苗生长和光合作用的影响. 应用生态学报, 2019, 30(5): 1445-1453.
21. 钱多多, 匡旭, 王绪高, 蔺菲, 原作强, 叶吉, 郝占庆. 2006—2017年长白山阔叶红松林木本植物种子雨的时空动态. 应用生态学报, 2019, 30(5): 1484-1493.
22. Shihong Jia, Xugao Wang, Zuoqiang Yuan, Fei Lin, Ji Ye, Zhanqing Hao, and Matthew Scott Luskin. 2018. Global signal of top-down control of terrestrial plant communities by herbivores. PNAS. 115(24): 6237-6242
23. Yuan Zuoqiang, Wang Shaopeng, Ali Arshad, Gazol Antonio, Ruiz-Benito Paloma, Wang Xugao, Lin Fei, Ye Ji, Hao Zhanqing and Loreau Michel. 2018. Aboveground carbon storage is driven by functional trait composition and stand structural attributes rather than biodiversity in temperate mixed forests recovering from disturbances. Annals of Forest Science. 75(3):67-79
24. Yunyun Wang, Robert P. Freckleton, Bojian Wang, Xu Kuang, Zhaochen Zhang, Zuoqiang Yuan, Fei Lin, Ji Ye, Xugao Wang, Zhanqing Hao. 2018. The role of breeding system in community dynamics: growth and mortality in forests of different successional stages. Ecology and Evolution. 2018: 1-12.
25. Zuoqiang Yuan, Arshad Ali, Shaopeng Wang, Antonio Gazol, Robert Freckleton, Xugao Wang, Fei Lin, Ji Ye, Li Zhou, Zhanqing Hao*, and Michel Loreau, 2018, Abiotic and biotic determinants of coarse woody productivity in temperate mixed forests. Science of The Total Environment. 630(51): 422–431
26. Xugao Wang, Thorsten Wiegand, Kristina J. Anderson-Teixeira, Norman A. Bourg, Zhanqing Hao, Robert Howe, Guangze Jin, David A. Orwig, Marko J. Spasojevic, Shunzhong Wang, Amy Wolf, Jonathan A. Myers. 2018. Ecological drivers of spatial community dissimilarity, species replacement, and species nestedness across temperate forests, Global Ecology and Biogeography. 1-12
27. Tan, Zheng-Hong; Hughes, Alice; Sato, Tamotsu; Zhang, Yi-Ping; Han, Shi-Jie; Kosugi, Yoshiko; Goulden, Michael; Deng, Xiao-Bao; Cao, Min; Hao, Zhan-Qing; Hu, Yue-Hua; Yu, Gui-Rui; Ma, Ke-Ping. 2017.
28. Quantifying forest net primary production: combining eddy flux, inventory and metabolic theory. IFOREST-BIOGEOSCIENCES AND FORESTRY, V(10) 475-482
29. Umana, Maria Natalia; Mi, Xiangcheng; Cao, Min; Enquist, Brian J.; Hao, Zhanqing; Howe, Robert; Iida, Yoshiko; Johnson, Daniel; Lin, Luxiang; Liu, Xiaojuan; Ma, Keping; Sun, I-Fang; Thompson, Jill. The role of functional uniqueness and spatial aggregation in explaining rarity in trees. GLOBAL ECOLOGY AND BIOGEOGRAPHY, 26(7), pp 777-786, 2017/7.
30. Jacob Usinowicz, Chia-Hao Chang-Yang, Yu-Yun Chen, James S. Clark, Christine Fletcher, Nancy C. Garwood, Zhanqing Hao, Jill Johnstone, Yiching Lin, Margaret R. Metz, Takashi Masaki, Tohru Nakashizuka, I-Fang Sun, Renato Valencia, Yunyun Wang, Jess K. Zimmerman, Anthony R. Ives & S. Joseph Wright, Temporal coexistence mechanisms contribute to the latitudinal gradient in forest diversity. Nature, 550(7674), pp 105-+, 2017/10/5.
31. Kuang, X., Zhu, K., Yuan, Z., Lin, F., Ye, J., Wang, X., Wang, Y., Hao, Z. 2017. Conspecific density dependence and community structure: Insights from 11 years of monitoring in an old-growth temperate forest in Northeast China.. Ecology and Evolution. 7(14), 5191–5200.
32. Zhang ZC, Papaik MJ, Wang XG, Hao ZQ, Ye J, Lin F and Yuan ZQ(2017)The effect of tree sizes, neighborhood competition and enviromant on tree growth in an old-growth temperate forest. Journal of Plant Ecology, 10: 970-980.
33. Yunyun Wang, Jian Zhang, JaleneM. LaMontagne, Fei Lin, Buhang Li, Ji Ye, Zuoqiang Yuan, XugaoWang & Zhanqing Hao (2017). Variation and synchrony of tree species mast seeding in an old-growth temperate forest. Journal of Vegetation Science. 28(2),413-423
34. 蔡礼蓉, 匡旭, 房帅, 原作强, 蔺菲, 叶吉, 郝占庆, 王绪高(2017)长白山阔叶红松林3个常见树种径向生长的影响因素. 应用生态学报, 28(5), 1407-1413.]
35. 米湘成, 郭静, 郝占庆, 谢宗强, 郭柯, 马克平, 中国森林生物多样性监测: 科学基础与执行计划. 生物多样性2016, 24 (11): 1203–1219
36. Wang XG, Wiegand T, Kraft NJB, Swenson NG, Davies SJ, Hao ZQ, Howe R, Lin Y, Ma KP, Mi XC, Su SH, Sun IF, Wolf A. (2016) Stochastic dilution effects weaken deterministic effects of niche-based processes in species rich forests. Ecology 97:347-360.
37. Yuan ZQ. Gazol A, Wang XG, Lin F, Ye J, Zhang Z, Suo YY, Kuang X, Wang YY, Jia SH, Hao ZQ.(2016) Pattern, dynamics and determinants of biomass stock in an old growth temperate forest: the role of habitat and tree size. Acta Oecologica 75: 15-23.
38. Suo Yanyan, Zuoqiang Yuan, Fei Lin, XugaoWang, Ji Ye,Edith Bai, Zhanqing Hao. (2016) Local-scale determinants of elemental stoichiometry of soil in an old growth forest. Plant Soil. 408:401–414
39. Yuan ZQ. Gazol A, Lin F, Wang XG, Ye J,Suo YY, Fang S, Mellard J, Hao ZQ.(2016). Scale-dependent effect of biotic interactions and environmental conditions in community assembly: insight from a large temperate forest plot. Plant Ecology. 217:1003-1014
40. Yuan ZQ, Wang SP, Gazol A, Mellard J, Lin F, Ye J, Hao ZQ, Wang XG, Loreau M. (2016) Multiple meterics of diversity have different effect s on termperate forest functioning over succession. Oeoloigca. 182(4): 1175-1185
41. Anderson-Teixeira, K. J., S. J. Davies, A. C. Bennett, E. B. Gonzalez-Akre, H. C. Muller-Landau, S. J. Wright, K. Abu Salim, A. M. A. Zambrano, A. Alonso, J. L. Baltzer, Y. Basset, N. A. Bourg, E. N. Broadbent, W. Y. Brockelman, S. Bunyavejchewin, D. Burslem, N. Butt, M. Cao, D. Cardenas, G. B. Chuyong, K. Clay, S. Cordell, H. S. Dattaraja, X. B. Deng, M. Detto, X. J. Du, A.
42. Duque, D. L. Erikson, C. E. N. Ewango, G. A. Fischer, C. Fletcher, R. B. Foster, C. P. Giardina, G. S. Gilbert, N. Gunatilleke, S. Gunatilleke, Z. Q. Hao, W. W. Hargrove, T. B. Hart, B. C. H. Hau, F. L. He, F. M. Hoffman, R. W. Howe, S. P. Hubbell, F. M. Inman-Narahari, P. A. Jansen, M. X. Jiang, D. J. Johnson, M. Kanzaki, A. R. Kassim, D. Kenfack, S. Kibet, M. F. Kinnaird, L. Korte, K. Kral, J. Kumar, A. J. Larson, Y. D. Li, X. K. Li, S. R. Liu, S. K. Y. Lum, J. A. Lutz, K. P. Ma, D. M. Maddalena, J. R. Makana, Y. Malhi, T. Marthews, R. M. Serudin, S. M. McMahon, W. J. McShea, H. R. Memiaghe, X. C. Mi, T. Mizuno, M. Morecroft, J. A. Myers, V. Novotny, A. A. de Oliveira, P. S. Ong, D. A. Orwig, R. Ostertag, J. den Ouden, G. G. Parker, R. P. Phillips, L. Sack, M. N. Sainge, W. G. Sang, K. Sri-ngernyuang, R. Sukumar, I. F. Sun, W. Sungpalee, H. S. Suresh, S. Tan, S. C. Thomas, D. W. Thomas, J. Thompson, B. L. Turner, M. Uriarte, R. Valencia, M. I. Vallejo, A. Vicentini, T. Vrska, X. H. Wang, X. G. Wang, G. Weiblen, A. Wolf, H. Xu, S. Yap, and J. Zimmerman. 2015. CTFS-ForestGEO:a worldwide network monitoring forests in an era of global change. Global Change Biology 21:528-549.
43. Li, H., X. G. Wang, C. Liang, Z. Q. Hao, L. S. Zhou, S. Ma, X. B. Li, S. Yang, F. Yao, and Y. Jiang. 2015. Aboveground-belowground biodiversity linkages differ in early and late successional temperate forests. Scientific Reports 5.
44. Pei, N. C., D. L. Erickson, B. F. Chen, X. J. Ge, X. C. Mi, N. G. Swenson, J. L. Zhang, F. A. Jones, C. L. Huang, W. H. Ye, Z. Q. Hao, C. F. Hsieh, S. Lum, N. A. Bourg, J. D. Parker, J. K. Zimmerman, W. J. McShea, I. C. Lopez, I. F. Sun, S. J. Davies, K. P. Ma, and W. J. Kress. 2015. Closely-related taxa influence woody species discrimination via DNA barcoding:evidence from global forest dynamics plots. Scientific Reports 5.
45. Wang, X. G., T. Wiegand, N. G. Swenson, A. T. Wolf, R. W. Howe, Z. Q. Hao, F. Lin, J. Ye, and Z. Q. Yuan. 2015. Mechanisms underlying local functional and phylogenetic beta diversity in two temperate forests. Ecology 96:1062-1073.
46. 张姗, 蔺菲, 原作强, 匡旭, 贾仕宏, 王芸芸, 索炎炎, 房帅, 王绪高, 叶吉, 郝占庆. 2015. 长白山阔叶红松林草本层物种多度分布格局及其季节动态. 生物多样性. 23(05):641-648.