姓名张宪省学历博士职称教授
所属部门植物学系
招生专业发育生物学（博士、硕士）
联系方式电话：** E-mail：zhangxs@sdau.edu.cn
::教师简介::
张宪省 博士、教授、博士生导师，山东农业大学副校长。现担任中国植物学会植物细胞生物学专业委员会副主任，中国遗传学会理事，山东省遗传学会理事长，山东省植物学会副理事长，中国作物学会作物生物技术分会理事，中国作物学会分子育种分会常务理事，美国植物生物学会会员。《Molecular Plant》、《JIPB》、《植物学报》等学术刊物编委。现为国家“973”项目首席科学家，享受国务院特殊津贴，入选国家“百千万”人才工程第一、二层次，入选山东省“泰山学者攀登计划”，山东省有突出贡献的中青年专家，山东省专业技术拔尖人才。国家实验教学示范中心联席会植物（动物）学科组组长、农业生物学国家级实验教学中心主任，生物化学与分子生物学国家级教学团队带头人。
::教学工作::
承担本科生《植物生命科学进展》；硕士研究生《植物发育分子生物学》、《植物发育生物学专题讨论》，博士研究生《生命科学进展》。另外，国家级实验教学示范中心全国植物、动物学科组组长，生物化学与分子生物学系列课程国家级教学团队建设项目负责人。
::研究方向::
1. 植物发育的分子基础: 本研究方向主要研究植物开花时间、花器官发育、授粉、种子发育等植物发育的重要科学问题，特别重视研究激素调节植物器官发育的分子机制。研究目的旨在解析上述发育过程的基因调控网络，阐明植物发育的细胞分子基础。
2. 植物基因工程: 本研究方向主要利用植物分子生物学研究的成果和基因工程技术，改良小麦和玉米的重要农艺性状和品质性状，提高作物产量，改良作物品质，产品直接服务于农业和工业生产。
::科研项目::
国家重大科学研究计划项目（973计划）：植物传粉与受精的分子调控网络（编号2013CB945100），2013～2017年，经费2900万元，项目首席科学家。 国家自然科学基金重点项目：植物激素调节体细胞胚胎发生的分子机理研究（编号**），2013～2015年，经费280万元，主持。 国家自然科学基金重点项目：植物激素调节体细胞胚胎发生的分子机理研究（编号**），2010～2013年，经费180万元，主持。高产转基因玉米新品种培育：转基因生物新品种培育重大专项课题（编号：2013ZX08003-003），2013年，经费100万元，共同主持。山东省农业生物资源创新利用研究课题：高产优质彩糯小麦种植材料创新与新品种培育，2013～2015年，经费100万元，主持。
::学术论文::
1. Su YH, Liu YB and Zhang XS (2011) Auxin-Cytokinin Interaction Regulates Meristem Development Molecular Plant doi: 10.1093/mp/ssr007.
2. Li X, Chen GH, Zhang WY, Zhang XS (2010) Genome-wide transcriptional analysis of maize endosperm in response to ae wx double mutation. JGG. 37: 749-762
3. Su YH, Cheng ZJ, Su YX, Zhang XS (2010) Pattern analysis of stem cell differentiation during in vitro Arabidopsis organogenesis. Front. Biol. 5(5): 464-470
4. Cheng ZJ, Zhu SS, Gao XQ, Zhang XS (2010) Cytokinin and auxin regulates WUS induction and inflorescence regeneration in in vitro in Arabidopsis. Plant Cell Rep 29:927-933.
5. Gao XQ, Zhu DZ, ZhangXS (2010) Stigma factors regulating self-compatible pollination. Front. Biol. 5(2): 156-163
6. Li XG, Su YH, Zhao XY, Li W, Gao XQ, Zhang XS (2010) Cytokinin overexpression-caused alteration of flower development is partially mediated by CUC2 and CUC3 in Arabidopsis. Gene 450:109-120
7. Guo J, Wang F, Song J, Zhang XS (2009) The expression of Orysa;CycB1;1 is essential for endosperm formation and causes embryo enlargement in rice. Planta 231:293-303
8. Ma Y, Wang F, Guo J, Zhang XS (2009) Rice OsAS2 Gene, a Member of LOB Domain Family, Functions in Regulation of Shoot Differentiation and Leaf Development. Journal of Plant Biology 52:374-381
9. Su YH, Zhao XY, Liu YB, Zhang CL, O’Neill SD, Zhang XS (2009) Auxin-induced WUS expression is essential for embryonic stem cell renewal during somatic embryogenesis in Arabidopsis. Plant Journal 59:448-460
10. Su YH and Zhang XS (2009) Auxin gradients trigger de novo formation of stem cells during somatic embryogenesis. Plant Siganling and Behavior 4:7, 574-576
11. Zhou Y, Zhang XJ, Kang XJ, Zhao XY, Zhang XS, Ni M (2009) SHORT HYPOCOTYL UNDER BLUE1 associates with MINISEED3 and HAIKU2 promotes in vivo to regulate Arabidopsis seed development. Plant Cell 21:106-117
12. Ma Q, Dai X, Xu Y, Guo J, Liu Y, Chen N, Xiao J, Zhang D, Xu Z, Zhang XS, Chong K. (2009) Enhanced tolerance to chilling stress in OsMYB3R-2 transgenic rice is mediated by alteration in cell cycle and ectopic expression of stress genes. Plant Physiology 150:257-271
13. Luo Y, Liu YB, Dong YX, Gao XQ, Zhang XS. (2009) Expression of a putative alfalfa helicase increases tolerance to abiotic stress in Arabiodopsis by enhancing the capacities for ROS scavenging and osmotic adjustment. Journal of Plant Physiology 166:385-394
14. Wang B, Sang Y, Song J, Gao XQ, Zhang XS. (2009) Expression of a rice OSARGOS gene in Arabidopsis promotes cell division and expension and increases organ size. J. Genet. Genomics 36:31-40
15. Sun Z, Ma L, Murphy RW, Zhang XS, Huang DW. (2009) Factors affecting mito-nuclear codon usage interactions in the OXPHO system of Drosophila melanogaster. J. Genet. Genomics 35:729-735
16. Sun Z, Wan DG, Murphy RW, Ma L, Zhang XS, Huang DW. (2009) Comparison of base composition and codon usage in insect mitochondrial genomes. Genes & Genomics 31:65-71
17. 于明明，李兴国，张宪省。APETALA1 启动子驱动AtPT4在转基因拟南芥中表达导致花和花器官发育异常。植物学报 44（1）：59－68。
18. Wu XQ, Li XG, Zhang XS. (2008) Molecular analysis of hormone-regulated pedal regeneration in Petunia. Plant Cell Reports 27:1169-1176
19. Zhao XY, Su YH, Cheng ZH, Zhang XS. (2008) Cell fate switch during in vitro plant organogenesis. Journal of Intergrative Plant Biology 50 (7):814-822
20. An BY, Luo Y, Li JR, Qiao WH, Zhang XS, Gao XQ. (2008) Expression of a vacuolar Na+/H+ antiportor gene of alfalfa enhances salinity tolerance in transgenic Arabidopsis. Acta Agronomica Sinica 34:557-564
21. Guo J, Song J, Wang F, Zhang XS. (2007) Genome-wide identification and expression analysis of rice cell cycle genes. Plant Molecular Biology 64:349-360
22. Qiao WH, Zhao XY, Li W, Zhang XS. (2007) Characterization of a root-specific vacuolar Na+/H+ antiporter gene in Agropyron elongatum and its expression involved in the salt tolerance of plants. Plant Cell Reports 26:1663-1672
23. Dong YX, Zhao XY, Wang JW, Yuan GL, Zhang XS. (2007) Improvement for Agronomic Traits of Partial Waxy Wheat by Combination of Backcrossing with a PCR-based DNA Marker. Jounal of Genet. Genomics 34:836-841
24. Guo SJ, Zhou HY, Zhang XS, Li XG, Meng QW. (2007) Overexpression of CaHSP26 in transgenic tobacco alleviates photoinhibition of PSII and PSI during chilling stress under low irradiance. Journal of Plant Physiology 164:126-136
25. Chu ZQ, Chen H, Zhang YY, Zhang ZH, Zheng NY, Yin BJ, Yan HY, Zhu L, Zhao XY, Yuan M, Zhang XS, and Xie Q. (2007) Knockout of the AtCESA2 gene affects microtubule orientation and causes abnormal cell expansion in Arabiodpsis thaliana. Plant Physiology143:213-224
26. Ma XJ, Tao Y, Zhao XY, Zhang XS. (2007) Wheat TaAS2, a member of LOB-Domain family, affects the adaxial-abaxial polarity of leaves in transgenic Arabidopsis. Plant Science 172: 181-188
27. Wang F, Huo SN, Guo J, Zhang XS. (2006) Wheat D-Type cyclin Triae;CYCD2;1 regulate development of transgenic Arabidopsis plants. Planta 224:1129-1140
28. Guan CM, Zhu SS, Li XG, Zhang XS. (2006) Inflorescence induction by hormones from Arabidopsis callus in vitro. Plant Cell Reports 25:1133-1137
29. Zhao W, Su HY, Song J, Zhao XY, Zhang XS. (2006) Ectopic expression of TaYAB1, a member of YABBY gene family in wheat, causes the partial abaxialization of the adaxial epidermises of leaves and arrests the development of shoot apical meristem in Arabidopsis, Plant Science 170:364-371
30. Guo SJ, Li JR, Qiao WH, Zhang XS. (2006) Analysis of amylose accumulation during seed development of maize. Acta Genetica Sinica 32:846-854
31. Zhao XY, Liu MS, Li JR, Guan CM, Zhang XS. (2005) The wheat TaGI1, involved in photoperoidic flowering, encodes an Arabidopsis GI ortholog. Plant Molecular Biology 58:53-64
32. Zhao XY, Cheng ZJ, Zhang XS. (2005) Overexpression of TaMADS1, a SEPALLATA-like gene in wheat, causes early flowering and the abnormal development of floral organs in Arabidopsis. Planta 223:698-707
33. Su HY, Li QZ, Li XG, Zhang XS. (2005) Characterization and expression analysis of a MADS box gene, HoMADS2, in Hyacinthus orientalis L. Acta Genetica Sinica 32: 1191-1198
34. Liu MS, Wang F, Dong YX, Zhang XS. (2005) Expression analysis of dihydroflavonol 4-reductase genes involved in anthocyanin biosynthesis in purple grains of Triticum aestivum. Journal of Intergrative Plant Biology 47 (9): 1107-1114
35. Li JR, Zhao W, Li QZ, Ye XG, An BY, Li X, Zhang XS. (2005) RNA Silencing of waxy gene results in low levels of amylose in the seeds of transgenic wheat (Triticum aestivum L.). Acta Genetica Sinica 32: 846-854
36. Xu HY, Li XG, Li QZ, Bai SN, Lu WL, Zhang XS. (2004) Characterization of HoMADS1 and its induction by plant hormones during in vitro development in Hyacinthus orientalis L. Plant Molecular Biology 55:209-220
37. Lan LF, Chen W, Lai Y, Suo JF, Kong ZS, Can Li, Lu Y, Zhang YJ, Zhao XY, Zhang XS, Zhang YS, Han B, Cheng J, Xue YB. (2004) Mornitoring of gene expression profiles and isolation of candidate genes involved in pollination and fertilization in rice (Oryza sativa L.) using 10K cDNA microarray. Plant Molecular Biology 54:471-483
38. An YR, Li XG, Su HY, Zhang XS. (2004) Pistil induction by hormones from callus of Oryza sativa in vitro. Plant Cell Reports 23:448-452
39. Li JR, Wang F, Zhao XY, Dong YX, Zhang LY, An BY, Zhang XS. (2004) Analysis of Seed Expressed Sequence Tags in Triticum aestivum. Acta Botanica Sinica 46 (3): 363-370
40. Zhang XS, Wang X, Li QZ, Huo SN, Li XG, Ye BX. (2003) A seed-specific gene RS6, encodes a putative cupin domain protein, is essential for seed development in Oryza sativa L. The Annual Meeting of the American Society of Plant Biologists, Plant Biology, Abstract p491
41. Li QZ, Li XG, Bai SN, Lu WL, Zhang XS. (2002) Isolation of HAG1 and its regulation by plant hormones during in vitro floral organogenesis in Hyacinthus orientalis L. Planta 215:533-540
42. Li QZ, Li XG, Bai SN, Lu WL, Zhang XS. (2001) Isolation and expression of HAP2 in Hyacinthus orientalis L. Development & Reproductive Biology10: 69-75
43. Zhang XS, Li QZ, Li XG, Bai SN, Lu WL. (2001) Molecular cloning and expression analysis of HAG1 in the floral organs of Hyacinthus orientalis L. Science in China C, 2000, 43: 395-401
44. Zhang XS, Li QZ, Li XG, Bai SN, Lu WL. (2000) Regulation of homeotic gene expression in the regenerated flowers of Hyacinthus. Plant Biology, The Annual Meeting of the American Society of Plant Physiologists 80: p39
45. Li QZ, Li XG, Sun JQ, Zhang XS. (2000) Isolation and expression of an AGAMOUS homolog in the flower of Cucumber (Cucumis sativas L.). Development & Reproductive Biology 9: 69-76
46. Zhang XS, Zheng CC, O'Neill SD. (1999) Expression of ethylene biosynthetic genes regulated by pollination-associated factors in Doritaenopsis flowers. Acta Botanica Sinica 41 (5): 468-471
47. Zheng CC, Zhang XS. (1999) Cloning and characterization of a circadian rhythm-regulated psaH cDNA in Pharbitis nil. Acta Botanica Sinica 41 (8): 820-824
48. Zhang XS, Li QZ, Zheng CC. (1999) Hormonal regulation of postpollination development of Doritaenopsis flowers by auxin and ethylene. Acta Phytophysiologica Sinica 25 (2): 178-186
49. Wang L, Zhang XS, Zhong HW, Li QZ. (1999) Expression and sequence analysis of a cDNA relative to Orchid ovule development. Acta Botanica Sinica 41(3): 276-279
50. Nadeau JA, Zhang XS, Li J, O'Neill SD. (1996) Ovule development: Identification of stage-specific and tissue-specific cDNAs. Plant Cell 8:213-239
51. O'Neill SD, Zhang XS, Zheng CC. (1994) Dark and circadian regulation of mRNA accumulation associated with photoperiodic flower induction. Plant Physiology104:569-580
52. Zhang XS, Nadeau JA, Li J, O'Neill SD. (1994) Isolation of cDNA clones specific to developing and mature ovules of orchid flowers. Plant Physiology 124: Supplement
53. Zhang XS, O'Neill SD. (1993) Ovary and gametophyte development are coordinately regulated by auxin and ethylene following pollination. Plant Cell 5:403-418
54. Nadeau JA, Zhang XS, Nair H, O'Neill SD. (1993) Temporal and spatial regulation of 1-aminocyclopropane-1-carboxylate oxidase in the pollination-induced senescence of orchid flowers. Plant Physiology 103:31-39
55. O'Neill SD, Nadeau JA, Zhang XS, Bui AQ, Hallev AH. (1993) Interorgan regulation of ethylene biosynthetic genes by pollination. Plant Cell 5:419-432
::科研奖励::
1.高等植物花发育的分子生物学研究, 1997, 山东省科技进步二等奖。
2. 小麦品质性状基因的分离与淀粉品质性状的遗传改良, 2005, 山东省自然科学二等奖。
::科研成果::
先后获得山东省自然科学奖和科技进步奖四项，申请基因发明专利六项，申请植物新品种保护权一项。先后在Plant Cell（10.4）、PNAS（9.5）、PLoS Genetics（9.5）等重要学术刊物发表高水平学术论文60余篇。指导本科生、硕士生、博士生的学位毕业论文有十余篇获得山东省优秀学位论文奖。
::研究专利::
1. 小麦周期蛋白基因及其分离方法与应用, 2002, 专利号**。
2. 偃麦草钠离子反向转运蛋白基因及其克隆与应用，2002，专利号**。
3. 紫色小麦二氢黄醇酮还原酶基因及其克隆与应用, 2005, 专利号**48。
4. 小麦TaGI1基因及其克隆与应用, 2005, 专利号**67。
5. 紫花苜蓿反向转运蛋白基因及其克隆与应用,2005,专利号**52。
6. 山农紫糯 1号，审核中，农业部新品种保护权。
