姓 名:
袁训来 性 别:
男
职 务:
现代古生物学和地层学国家重点实验室主任 职 称:
研究员
通讯地址:
南京市北京东路39号
邮政编码:
210008 电子邮件:
xlyuan@nigpas.ac.cn


简历：
1983-1987：北京大学地质系，地层学古生物学专业，获学士学位
1987-1990：北京大学地质系，地层学古生物学专业，获硕士学位
1990-1993：中国科学院南京地质古生物研究所实习研究员
1993-1996：中国科学院南京地质古生物研究所助理研究员
1997-1999：中国科学院南京地质古生物研究所副研究员
2000-现今：中国科学院南京地质古生物研究所研究员
1999-2000：Organismic & Evol. Biology, Harvard University，访问****
1995-1996：Department of Geology, University Of Montreal，访问****


研究领域：
　　多细胞生物起源和早期演化。

社会任职：

获奖及荣誉：
国家****基金（2006）、第八届江苏省青年科技奖（2003）、美国古生物学杂志特别论文奖（2002）、中国地质学会“银锤”奖（2003）、第六届南京市十大科技之星（2005），第五届中国古生物学会尹赞勋奖（2004）、首批“新世纪百千****才工程”国家级人选（2004）、江苏省“333工程”人选、全国优秀科技工作者（2014）；2005年度科学中国人、首届中科院研究生院优秀教师（2006）、第三届中国科学院创新文化建设先进个人（2007）、江苏省科普先进个人等（2003）；专著《陡山沱期生物群》获中国科技图书一等奖、安徽省图书一等奖和中国图书奖（2003）；国务院特殊津贴（2007）；“蓝田生物群”研究获得2013年度江苏省科技成果一等奖。2014年，第六届全国优秀科技工作者；2015中国科学院朱李月华奖。
在贵州瓮安地区发现的似地衣化石被评为2005年度“中国基础研究十大新闻”；在湖北三峡地区发现的6.3亿年前的动物胚胎化石被评为2007年度“中国基础研究十大新闻”；“蓝田生物群”研究被评为2011年度中国地质科学十大进展等。


承担科研项目情况：
1. 国家自然科学基金“新元古代陡山沱期后生植物多样性及动物化石的研究”（2000.01-2002.12）
2. 中国科学院知识创新重大方向性项目“新元古代雪球事件前后生物演化及环境背景”（2000-2003）
3. 973科技部国家重点基础研究“重大地史时期生物的起源、辐射、灭绝和复苏”子课题“多细胞生物的早期演化、后生动物的起源及其环境条件”（2000-2005）
4. 科技部国际科技合作重点项目“多细胞生物的早期演化、后生动物的起源及其环境条件”（2003-2008）
5. 973科技部国家重点基础研究“重大地史时期生物的起源、辐射、灭绝和复苏”子课题“多细胞生物的早期演化、后生动物的起源及其环境条件”（2006-2010）
6. 国家****基金“元古代真核生物的演化”（2007.01-2010.12）
7. 国家自然科学基金“山西五台地区太古代与元古代过渡地层微体化石研究”（2005.01-2007.12）
8. 中国科学院知识创新重要方向性项目“华北地台晚太古代至古元古代化石生物学和古环境探索”（2009-2011）
9. 中石化前瞻性基础研究课题“中国新元古代海相地层研究”（2007-2009）
10. 中国科学院、国家外国专家局创新团队国际合作伙伴计划《关键地史时期生态系统的演变、崩溃和重建》子课题“新元古代大冰期前后生态系统的崩溃和重建”（2007-2012）
11. 国家基金委重点项目“埃迪卡拉纪蓝田生物群研究”（2012.01-2016.12）
12. 中国科学院重点部署项目“多细胞生物起源与早期演化”（2011-2015）
13. 科技部973项目“从雪球事件到寒武纪大爆发：距今6亿年前后的生物与环境演变”子课题“多细胞生物起源和早期演化”（2013.01-2017.12）
14. 中科院项目“蓝田生物群”研究（2013-2015）
15. 中科院前沿重点项目，“三峡埃迪卡拉生物群”研究（2016-2020）
16. 中科院专项B“关键地史时期生物与环境演变过程及其机制”子课题“埃迪卡拉纪真核生物的辐射和环境”（2019-2023）
17. 基金委创新群体“多细胞生物起源和早期演化”（2020-2024）

代表论著：
1.Zhang Yun and Yuan Xunlai, 1992: New data on multicellular thallophyte and fragments of cellular tissues from Late Proterozoic phosphate rocks, South China. Lethaia, 25:1-18. Pl.10, Appendix 1.
2.袁训来，王启飞，张昀，1993. 贵州瓮安磷矿晚前寒武纪陡山沱期的藻类化石群. 微体古生物学报，10（4），409-420.
3.陈孟莪，萧宗正，袁训来，1994. 晚震旦世的特种生物群落--庙河生物群新知. 古生物学报，33（40），391-403.
4.袁训来，李军，陈孟莪，1995. 晚前寒武纪后生植物的发展及其化石证据. 古生物学报，34（1），90-102.
5.Chen Menge, Xiao Zongzheng and Yuan Xunlai, 1995. A great diversification of macroscopic algae in Neoproterozoic. Scientia Geologica, 4(3), 295-308.
6.张昀，袁训来，1995. 元古宙末多细胞红藻有性生殖结构的发现. 中国科学（B辑），25（7），749-754.
7.尹磊明，袁训来，1996. 中国南方震旦纪--早寒武世含磷地层中的微体浮游植物化石. 叶连俊主编《生物有机质成矿作用》，63-72. 海洋出版社，北京.
8.Yin Leiming and Yuan Xunlai, 1996: Neoproterozoic fossil records of algae and organic-walled microphytoplankton in China, and their implications. In: Pang Z., Zhang J., and Sun J., (eds.) Advances in solid Earth sciences. 132-136. Science Press, Beijing.
9.Yin Leiming and Yuan Xunlai, 1996. Microphytoplanktonic fossils of the Sinian-Early Cambrian phosphorite deposits in the Yangtze Plateform. In Ye L. (ed.) Aspects of Biomineralization, 51-65. Seismological Press, Beijing.
10.Zhang Yun and Yuan Xunlai, 1996. Sexual reproductive structures of Latest Proterozoic multicellular rhodophytes (red algae) from South China. Science in China (Series C), 39(1), 28-39.
11.Yuan Xunlai and Hofmann H. J., 1998. New microfossils from the Neoproterozoic (Sinian) Doushantuo Formation, Wengan, Guizhou Province, southwestern China. Alcheringa, 22(3), 189-222.
12.Xiao Shuhai, Knoll A. H. and Yuan Xunlai, 1998. Morphological reconstruction of Miaohephyton bifurcatum, a possible brown alga from the Neoproterozoic Doushantuo Formation, South China. Journal of Paleontology, 72(6), 1072-1086.
13.周传明，袁训来，薛耀松，1998. 贵州瓮安磷矿陡山沱期假化石的研究. 《微体古生物学报》，15（4），380-384.
14.Zhang Yun, Yuan Xunlai and Yin Leiming, 1998. Interpreting Late Precambrian Microfossils. Science, 282(5395), 1783.
15.袁训来，1999. 新元古代陡山沱期瓮安生物群研究简况. 《微体古生物学报》，16（3），281-286.
16.袁训来，周传明，1999. 瓮安生物群. 江苏省科学技术协会编，科技进步与学科发展—迎接21世纪挑战. 301-304页.
17.Yuan Xunlai, Li Jun and Cao Ruiji, 1999. A diverse metaphyte assemblage from the Neoproterozoic black shales of South China. Lethaia, 32, 143-155.
18.袁训来，周传明，1999. 新元古代磷酸盐化微化石. 江苏地质，23（4），202-208.
19.Xiao Shuhai, Yuan Xunlai, Knoll A. H., 2000. Eumetazoan Fossils in Terminal Proterozoic Phosphorites? PNAS, 97(25), 13684-13689.
20.Yuan Xunlai, Xiao Shuhai, Li Jun, Yin Leiming, and Cao Ruiji, 2001. Pyritized chuarids with excystment structures from the late Neoproterozoic Lantian Formation in Anhui, South China. Precambrian Research, 107(3-4), 253-263.
21.周传明，燕夔，胡杰，肖书海，鲍惠铭，袁训来等，2001. 皖南新元古代两期冰期事件. 地层学杂志，25（4），247-258.
22.袁训来，2000. 地球生命起源的假说. 徐均涛主编，中学生探索丛书—《进化长河-古生物》. 江苏科学技术出版社，1-6页.
23.袁训来，2000. 地球曾经是一个“雪球”吗？徐均涛主编，中学生探索丛书—《进化长河-古生物》. 江苏科学技术出版社，12-16页.
24.尹磊明，曹瑞骥，袁训来，王宗哲，2001. 上前寒武系. 周志毅主编，《塔里木盆地各纪地层》，科学出版社，1-11.
25.林焕令，陈挺恩，袁金良，袁文伟，张允白，袁训来，章森桂，2001. 寒武系. 周志毅主编，《塔里木盆地各纪地层》，科学出版社，12-38.
26.尹磊明，肖书海，袁训来，2001. 对贵州瓮安陡山沱磷块岩中针状结构物的新认识. 科学通报，46(12)，1031-1036.
27.曹瑞骥，袁训来，肖书海，2001. 论锥叠层石群(Conophyton)的形态发生──对苏北新元古代九顶山组一个似锥叠层石的剖析. 古生物学报，40（3），318-329.
28.Xiao Shuhai, Yuan Xunlai, Steiner M., and Knoll A. H., 2002. Macroscopic carbonaceous compressions in a terminal Proterozoic shale: A systematic reassessment of the Miaohe Biota, South China. Journal of Paleontology, 76(2), 347-376.
29.袁训来，肖书海，尹磊明，Knoll A. H.，周传明，穆西南，2002. 陡山沱期生物群：早期动物辐射前夕的生命. 171页，中国科技大学出版社，合肥.
30.Zhou Chuanming, Yuan Xunlai, Xiao Shuhai, 2002. Phosphatized biotas from the Neoproterozoic Doushantuo Formation on the Yangtze Platform. Chinese Science Bulletin, 47(22), 1918-1924.
31.Yuan Xunlai, Xiao Shuhai, Parsley R. L., Zhou Chuanming, Chen Zhe, Hu Jie, 2002. Towering sponges in an Early Cambrian Lagersta¨tte: Disparity between nonbilaterian and bilaterian epifaunal tierers at the Neoproterozoic-Cambrian transition. Geology, 30(4), 363-366.
32.袁训来，姚锦仙，2002. 性起源、性分化和地球早期生物的多样性. 《科学》，54（5），24-26.
33.Xiao Shuhai, Hu Jie, Yuan Xunlai, Parsley R. L., and Cao Ruiji, 2005. Articulated sponges from the Lower Cambrian Hetang Formation in southern Anhui, South China: Their age and implications for the early evolution of sponges. Palaeogeography, Palaeoclimatology, Palaeoecology, 220, 89-117.
34.Xiao Shuhai, Bao Huiming, Wang Haifeng, Kaufman A. J., Zhou Chuanming, Li Guoxiang, Yuan Xunlai, Ling Hongfei, 2004, The Neoproterozoic Quruqtagh Group in eastern Chinese Tianshan: evidence for a post-Marinoan glaciation. Precambrian Research, 130, 1-26.
35.尹磊明，边立曾，袁训来，2003. 山西中元古代汝阳群分枝管状藻体和显微环螺旋加厚管体的发现. 中国科学（D），33（8），769-774.
36.Xiao Shuhai, Knoll A. H., Yuan Xunlai and Pueschel C. M., 2004. Phosphatized multicellular algae in the Neoproterozoic Doushantuo Formation, China, and the early evolution of florideophyte red algae. American Journal of Botany, 91, 214-227.
37.Cao Ruiji, Yuan Xunlai and Mu Xi’nan, 2003. Brief History and Current Status of Stromatolite Study in China. Acta Micropalaeontologica Sinica, 19(4), 5-14.
38.Yin Leiming, Yuan Xunlai, 2003. A review of eukaryotic fossils in the late Mesoproterozoic Ruyang Group in Shanxi, Chin. Acta Micropalaeontologica Sinica, 19(4), 39-46.
39.Mu Xi’nan, Yuan Xunlai, 2003. Introduction of Fossil algae and Stromatolites. Acta Micropalaeontologica Sinica, 19(4), 1-4.
40.Bao Huiming, Thiemens M. H., Loope D. B., Yuan Xunlai, 2003. Sulfate oxygen-17 anomaly in an Oligocene ash bed in mid-North America: Was it the dry fogs? Geophysical Research Letters, 30(16), 1843.
41.Zhou Chuanming, Tucker R., Xiao Shuhai, Peng Zhanxiong, Yuan Xunlai, Chen Zhe, 2004. New constraints on the ages of Neoproterozoic glaciations in south China. Geology, 32(5): 437-440.
42.Cao Ruiji and Yuan Xunlai, 2003. Pre-Sinian biostratigraphy of China. In: Zhang et al. eds. Biostratigraph of China. 1-30. Science Press, Beijing.
43.Yin Leiming and Yuan Xunlai, 2003. Sinian biostratigraphy of China. In: Zhang et al. eds. Biostratigraphy of China. 31-54. Science Press, Beijing.
44.Chen Zhe, Hu Jie, Zhou Chuanming, Xiao Shuhai, Yuan Xunlai, 2004. Sponge assemblage from early Cambrian Hetang Formation, southern Anhui. Chinese Science Bulletin, 49(15), 1625-1628.
45.Yin Leiming, Bian Lizeng, Yuan Xunlai, 2004. Discovery of branched tubular algae and microscopic tubes with annular-helical thickening from the Mesoproterozoic Ruyang Group of Shanxi, North China. Science in China (Ser. D), 47(10), 880-885.
46.Hua Hong, Chen Zhe, Yuan Xunlai, Xiao Shuhai, Zhang Luyi, 2005. Skeletogenesis and asexual reproduction in the earliest biomineralizing animal Cloudina. Geology, 33(4), 277-280.
47.周传明，袁训来，肖书海，陈哲，薛耀松，2004. 湖北保康陡山沱磷酸盐化微体化石组合. 微体古生物学报，21（4），349-366.
48.Yao Jinxian, Xiao Shuhai, Yin Leiming, Li Guoxiang and Yuan Xunlai, 2005. Basal Cambrian microfossils from the Yurtus and Xishanblaq formations (Tarim, north-west China): Systematic revision and biostratigraphic correlation of Micrhystridium-like acritarchs from China. Paleontology, 48(4), 687-708.
49.Xiao Shuhai, Hu Jie, Yuan Xunlai, Parsley R. L., Cao Ruiji, 2005. Articulated sponges from the Lower Cambrian Hetang Formation in southern Anhui, South China: their age and implications for the early evolution of sponges. Palaeogeography, Palaeoclimatology, Palaeoecology, 220, 89-117。
50.周忠和，骆永明，董云社，袁训来，高星，2005. 浅谈我国地球环境与生命过程研究中的一些科学问题. 科学通报，50（2），195-200.
51.Xiao Shuhai，Shen Bing，Zhou Chuanming， Xie Guwei， Yuan Xunlai，2005. A uniquely preserved Ediacaran fossil with direct evidence for a quilted bodyplan. PNAS, 102(29), 10227-10232.
52.Yuan Xunlai, Xiao Shuhai, Taylor T. N., 2005. Lichen-like symbiosis 600 million years ago. Science, 308, 1017-1020.
53.Meng Fanwei, Zhou Chuanming, Chen Zhilin, Yuan Xunlai, 2005. The oldest known dinoflagellates: morphological and molecular evidence from Mesoproterozoic rocks at Yongji, Shanxi Province. Chinese Science Bulletin, 50(12), 1230-1234.
54.Yin Leiming, Yuan Xunlai, Meng Fanwei and Hu Jie, 2005. Protists of the upper Mesoproterozoic Ruyang Group in Shanxi Province, China. Precambrian Research, 141, 29-66.
55.Hagadorn1J. W., Xiao Shuhai, Donoghue P. C. J., Bengtson S., Gostling N. J., Pawlowska M., Raff E. C., Raff R. A., Turner F. R., Yin Chongyu, Zhou Chuanming, Yuan Xunlai, McFeely M. B., Stampanoni M., Nealson K. H., 2006. Cellular and Subcellular Structure of Neoproterozoic Animal Embryos. Science, 314, 291-294.
56.Xiao Shuhai, Hagadorn J. W., Zhou Chauming, Yuan Xunlai, 2007. Rare helical spheroidal fossils from the Neoproterozoic Doushantuo lagerst?tte: Ediacaran animal embryos come of age? Geology, 35, 115-118.
57.袁训来，肖书海，周传明，2006. 新元古代陡山沱期真核生物的辐射. 戎嘉余（主编）：《生物的起源、辐射与多样性演变-华夏化石记录的启示》. 北京，科学出版社，13-27.
58.曹瑞骥，袁训来，2006. 《叠层石》. 合肥，中国科技大学出版社. 383页.
59.Yin Leiming, Zhu Maoyan, Knoll A. H., Yuan Xunlai, Zhang Junming and Hu Jie，2007. Doushantuo embryos preserved inside diapause egg cysts. Nature, 445, 198-201
60.Xiao Shuhai, Zhou Chuanming, Yuan Xunlai, 2007. Undressing and redressing Ediacaran embryos. Nature 445, 198-201.
61.Zhou Chuanming, Xie Guwei, Mcfadden K., Xiao Shuhai and Yuan Xunlai, 2007. The diversification and extinction of Doushantuo-Pertatataka acritarchs in South China: causes and biostratigraphic significance. Geological Journal, 42, 229-262.
62.Hua Hong, Chen Zhe and Yuan Xunlai, 2007. The advent of mineralized skeletons in Neoproterozoic Metazoa—new fossil evidence from the Gaojiashan Fauna. Geological Journal, 42, 263-279.
63.Dong Lin, Xiao Shuhai, Shen Bing, Yuan Xunlai, Yan Xianqin, Peng Yongbo, 2008. Restudy of the worm-like carbonaceous compression fossils Protoarenicola, Pararenicola, and Sinosabellidites from early Neoproterozoic successions in North China. Palaeogeography, Palaeoclimatology, Palaeoecology, 258, 138-161.
64.Yin Leiming, Yuan Xunlai, 2007. Radiation of Meso-Neoproterozoic and early Cambrian protists inferred from the microfossil record of China. Palaeogeography, Palaeoclimatology, Palaeoecology, 254, 350-361.
65.James D. S., Yin Leiming, Robert J. B., Kaufman A. J., Meng Fanwei, Hu Jie, Shen Bing, Yuan Xunlai, Bao Huiming and Xiao Shuhai, 2007. Ultrastructural and Geochemical Characterization of Archean–Paleoproterozoic Graphite Particles: Implications for Recognizing Traces of Life in Highly Metamorphosed Rocks. Astrobiology, 7(4), 684-704.
66.严贤勤，孟凡巍，袁训来，2006. 徐淮地区新元古代九顶山组燧石结核的地球化学特征. 微体古生物学报，23（3），295-302.
67.袁训来，2007. 话说六亿年前的动物胚胎化石. 生物进化，1，16-19.
68.彭永波，王丹，袁训来，2007. 长城系串岭沟组球形疑源类的超微结构. 微体古生物学报，24（2），194-204.
69.孟凡巍，袁训来等，2007. 同步辐射X射线荧光微探针技术测定太古宇条带状硅铁建造中硅质条待鸡包裹体的微量元素. 岩石学报，23（09），2079-2084.
70.Kathleen A. M., Huang Jing, Chu Xuelei, Jiang Ganqing, Kaufman A. J., Zhou Chuanming, Yuan Xunlai and Xiao Shuhai, 2008. Pulsed oxidation and biological evolution in the Ediacaran Doushantuo Formation. PNAS, 105(9), 3197-3202.
71.尹磊明，周传明，袁训来，2008. 湖北宜昌埃迪卡拉系陡山沱组天柱山乱囊胞—Tianzhushania的新认识. 古生物学报，47（2），129-140.
72.Peng Yongbo, Bao Huiming, Yuan Xunlai, 2009. New morphological observations for Paleoproterozoic acritarchs from the Chuanlinggou Formation, North China. Precambrian Research, 168, 223-232.
73.Xu Bei, Xiao Shuhai, Zou Haibo, Chen Yan, Li Zhengxiang , Song Biao, Liu Dunyi, Zhou Chuanming, Yuan Xunlai, 2009. SHRIMP zircon U–Pb age constraints on Neoproterozoic Quruqtagh diamictites in NW China. Precambrian Research, 168, 247-258.
74.袁训来，王丹，肖书海，2009. 新元古代陡山沱期的动物. 古生物学报，48(3)，375-389.
75.曹瑞骥，袁训来，2009. 中国叠层石研究进展. 古生物学报，48(3)，314-321.
76.Shen Bing, Xiao Shuhai, Zhou Chuanming, Yuan Xunlai, 2009. Yangtziramulus zhangi new genus and species, a carbonate-hosted Ediacaran macrofossil from the Dengying Formation in the Yangtze Gorges area, South China. Journal of Paleontology, 83, 575-587.
77.Shen Bing, Xiao Shuhai, Zhou Chuanming, Kaufman A. J., Yuan Xunlai, 2010. Carbon and sulfur isotope chemostratigraphy of the Neoproterozoic Quanji Group of the Chaidam Basin, NW China: Basin stratification in the aftermath of an Ediacaran glaciation postdating the Shuram event? Precambrian Research, 177, 241-252.
78.Zhou Chuanming, Bao Huiming, Peng Yongbo, Yuan Xunlai, 2010. Timing the deposition of 17O-depleted barite at the aftermath of Nantuo glacial meltdown in South China. Geology, 38, 903-906.
79.华洪，陈哲，袁训来，肖书海，蔡耀平，2010. 陕南伊迪卡拉纪末期的瓶状化石—可能最早的有孔虫化石. 中国科学，D辑，40（9），1105-1114.
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